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doi: 10.1186/1741-7015-9-91 7 BakkenIJTveitoKGunnesNGhaderiSStoltenbergCTrogstadLet al

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(ii) GxG (Gene x Gene) interaction, wherein the loss of two genes leads to a fitness phenotype that is not the simple additive sum of two single KOs, and (iii) GxGxE (Gene x Gene x Environment) interaction, wherein the impact of the pairwise genetic interaction is dependent on the metabolic state